The Cross-banded or Tawny Tree Frog is a drab frog with tan to light brown coloration, expanded disks on toes and fingers, and minimal webbing in the feet. They look very similar to the Masked Tree Frog, which is another species in the Smilisca genus. In Costa Rica, it is mainly found in the Caribbean lowlands. It has nocturnal habits and prefers to be on trees, however during the reproductive season, the males descend to the ground and emit their call from the edge of small ponds. Unlike other nocturnal species that I have photographed, this frog would never fully open the eyes. Its position suggests how sleepy they are during the day. They probably rely on camouflage rather than detecting predators and escaping from them.

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